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In the canonical pathway, the type I receptor phosphorylates the receptor-activated Smad proteins (R-Smads), Smad2 and Smad3 (Xu et al., 2018), allowing for the formation of heterotrimers with Smad4, and binding to the Smad binding element (SBE) (Zawel et al., 1998)

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[DOI] [PMC free article] [PubMed] [Google Scholar] 32.Curi T.C.P., de Melo M.P., de Azevedo R.B., Curi R